A transporter mishap converts several crewmembers into spy-themed holosuite characters, with their true memories erased and their lives linked directly to the program.
Paranoia swarms as Sisko and Odo work to prevent a military power-grab at Starfleet Command, while the Defiant is believed to have been overtaken by Changelings.
Kira assists Gul Dukat, now in command of a freighter, to stop a Klingon Bird-of-Prey that destroyed an outpost with a joint Bajoran and Cardassian delegation.
Miles struggles re-adjusting to normal life after having the realistic memories of serving a long, difficult prison sentence transplanted into his mind.
